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Flying fish


Flying fish

Flying fish
Temporal range: Miocene–Recent
Sailfin flying-fish
Parexocoetus brachypterus
Scientific classification
Kingdom: Animalia
Phylum: Chordata
Class: Actinopterygii
Order: Beloniformes
Family: Exocoetidae

The Exocoetidae are a family of marine fish in the order Beloniformes of class (biology) Actinopterygii. Fish of this family are known as flying fish. About 64 species are grouped in seven to nine genera. Flying fish can make powerful, self-propelled leaps out of water into air, where their long, wing-like fins enable gliding flight for considerable distances above the water's surface. This uncommon ability is a natural defense mechanism to evade predators.

The oldest known fossil of a flying or gliding fish, Potanichthys xingyiensis, dates back to the Middle Triassic, 235–242 million years ago. However, this fossil is not related to modern flying fish, which evolved independently about 66 million years ago.[1][2]


  • Etymology 1
  • Distribution and description 2
    • Flight measurements 2.1
  • Fishery and cuisine 3
  • Importance 4
    • Barbados 4.1
      • Maritime disputes 4.1.1
  • See also 5
  • References 6
  • External links 7


The term Exocoetidae is not only the present scientific name for a genus of flying fish in this family, but also the general name in Latin for a flying fish. The suffix -idae, common for indicating a family, follows the root of the Latin word exocoetus, a transliteration of the Ancient Greek name ἐξώκοιτος. This means literally "sleeping outside", from ἔξω "outside" and κοῖτος "bed", "resting place",[3] so named as flying fish were believed to leave the water to sleep on the shore.[4]

Distribution and description

Flying fish
Flying fish taking off

Flying fish live in all of the

  • May 2008 video of 45 second flight
  • Explanation
  • Flying Fish, National Geographic Society

External links

  1. ^ Oldest flying fish fossil found in China Nature, News, 31 October 2012.
  2. ^ Xu, Guang-Hui; Li-Jun Zhao, Ke-Qin Gao and Fei-Xiang Wu (2012) A new stem-neopterygian fish from the Middle Triassic of China shows the earliest over-water gliding strategy of the vertebrates" Proceedings of the Royal Society B: Biological Sciences, Published online before print October 31, 2012. doi:10.1098/rspb.2012.2261
  3. ^ Harper, Douglas. "exocet".  
  4. ^ Pliny's Natural History, vol. IX, 19)
  5. ^ Fish, F. E. (1990). Wing Design And Scaling of Flying Fish With Regard To Flight Performance. Journal of Zoology, 221(3), 391-403.
  6. ^ a b Davenport, J. (1994). How And Why Do Flying Fish Fly?. Reviews in Fish Biology and Fisheries, 4(2), 184-214.
  7. ^ Dasilao, J., & Yamaoka, K. (1998). Development of the Vertebral Column And Caudal Complex in a Flyingfish,Parexocoetus Mento Mento (Teleostei: Exocoetidae). Ichthyological Research, 45(3), 303-308.
  8. ^ a b Dasilao, J., & Yamaoka, K. (1998). Development Of The Vertebral Column And Caudal Complex In A Flyingfish,Parexocoetus Mento Mento (Teleostei: Exocoetidae). Ichthyological Research, 45(3), 303-308.
  9. ^ a b Dasilao, J. C., & Sasaki, K. (1998). Phylogeny Of The Flyingfish Family Exocoetidae (Teleostei, Beloniformes). Ichthyological Research, 45(4), 347-353.
  10. ^ a b Fish, F. E. (1990). "Wing design and scaling of flying fish with regard to flight performance" ( 
  11. ^ a b c d e f g Kutschera, U. (2005). "Predator-driven macroevolution in flyingfishes inferred from behavioural studies: historical controversies and a hypothesis" ( 
  12. ^ a b Fish, F. (1991). "On a fin and a prayer" ( 
  13. ^ a b "Fast flying fish glides by ferry".  
  14. ^ a b Ross Piper (2007), Extraordinary Animals: An Encyclopedia of Curious and Unusual Animals, Greenwood Press.
  15. ^ ExocoetidaeFlying Fish, National Geographic. Retrieved 10 August 2014.
  16. ^ Joseph Banks (1997). The Endeavour Journal of Sir Joseph Banks 1768–1771 (PDF).  
  17. ^ "Barbados/Trinidad and Tobago".  
  18. ^ "Claims of Caribbean piracy as national symbol takes flight". The Sydney Morning Herald. December 18, 2004. 


See also

In recent times, flying fish have also been gaining in popularity in other islands, fueling several maritime disputes. In 2006, the council of the United Nations Convention on the Law of the Sea[17] fixed the maritime boundaries between Barbados and Trinidad and Tobago over the flying fish dispute, which gradually raised tensions between the neighbours.[18] The ruling stated both countries must preserve stocks for the future. Barbadian fishers still follow the flying fish southward. Flying fish remain an important part of Barbados' main national dish.

Maritime disputes

Many aspects of Barbadian culture center around the flying fish: it is depicted on coins, as sculptures in fountains, in artwork, and as part of the official logo of the Barbados Tourism Authority. Additionally, the Barbadian coat of arms features a pelican and dolphin fish on either side of the shield, but the dolphin resembles a flying fish. Furthermore, actual artistic renditions and holograms of the flying fish are also present within the Barbadian passport.

Just after the completion of the Bridgetown Harbor / Deep Water Harbor in Bridgetown, Barbados saw an increase of ship visits, linking the island to the world. The overall health of the coral reefs surrounding Barbados suffered due to ship-based pollution. Additionally, Barbadian overfishing pushed them closer to the Orinoco delta, no longer returning to Barbados in large numbers. Today, the flying fish only migrate as far north as Tobago, around 120 nmi (220 km; 140 mi) southwest of Barbados. Despite the change, flying fish remain a coveted delicacy.

Barbados is known as "the land of the flying fish", and the fish is one of the national symbols of the country. Once abundant, it migrated between the warm, coral-filled Atlantic Ocean surrounding the island of Barbados and the plankton-rich outflows of the Orinoco River in Venezuela.



In the Solomon Islands, the fish are caught while they are flying, using nets held from outrigger canoes. They are attracted to the light of torches. Fishing is done only when no moonlight is available.

Flying fish are commercially fished in Japan, Vietnam, and China by gillnetting, and in Indonesia and India by dipnetting.[11] Often in Japanese cuisine, the fish is preserved by drying. The roe of Cheilopogon agoo, or Japanese flying fish, is used to make some types of sushi, and is known as tobiko. It is also a staple in the diet of the Tao people of Orchid Island, Taiwan. Flying fish is part of the national dish of Barbados, cou-cou and flying fish.

Fishery and cuisine

The flights of flying fish are typically around 50 meters (160 ft),[14] though they can use updrafts at the leading edge of waves to cover distances of up to 400 m (1,300 ft).[14][15] They can travel at speeds of more than 70 km/h (43 mph).[11] Maximum altitude is 6 m (20 ft) above the surface of the sea.[12] Some accounts have them landing on ships' decks.[11][16]

In May 2008, a Japanese television crew (NHK) filmed a flying fish (dubbed "Icarfish") off the coast of Yakushima Island, Japan. The fish spent 45 seconds in flight.[13] The previous record was 42 seconds.[13]

Flight measurements

Exocoetidae feed mainly on plankton. Predators include dolphins, tuna, marlin, birds, squids, and porpoises.[11]

Genus Exocoetus has one pair of fins and a streamlined body to optimize for speed, while Cypselurus has a flattened body and two pairs of fins, which maximize its time in the air. From 1900 to the 1930s, flying fish were studied as possible models used to develop airplanes.[11]

Research has shown that the flying fish has undergone morphological changes throughout its history, the first of which is fully broadened neural arches. Neural arches act as insertion sites for muscles, connective tissues, and ligaments in a fish’s skeleton. Fully broadened neural arches act as more stable and sturdier sites for these connections, creating a strong link between the vertebral column and cranium.[8] This ultimately allows a rigid and sturdy vertebral column (body) that is beneficial in flight. Having a rigid body during glided flight gives the flying fish aerodynamic advantages, increasing its speed and improving its aim.[8] Furthermore, flying fish have developed vertebral columns and ossified caudal complexes.[9] These features provide the majority of strength to the flying fish, allowing them to physically lift their body out of water and glide remarkable distances. These additions also reduce the flexibility of the flying fish, allowing them to perform powerful leaps without weakening midair.[9] At the end of a glide, it folds its pectoral fins to re-enter the sea, or drops its tail into the water to push against the water to lift itself for another glide, possibly changing direction.[10][11] The curved profile of the "wing" is comparable to the aerodynamic shape of a bird wing.[12] The fish is able to increase its time in the air by flying straight into or at an angle to the direction of updrafts created by a combination of air and ocean currents.[10][11]

As a result, flying fish have undergone natural selection in which species gain unique traits to better adapt to their environments. By becoming airborne, flying fish outsmart their predators and environment. This increase of speed and maneuverability is a direct advantage to flying fish, and has given them leverage when compared to other species in their environment. [7] Along with relationship difficulties, abiotic factors also play a part. Harsh ocean currents make it extremely difficult for small fish to survive in this habitat. Research suggests that difficult environmental factors in the flying fish's habitat have led to the evolution of modified fins.[6]

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