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Rhamnaceae

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Title: Rhamnaceae  
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Subject: Flora of Madhya Pradesh, List of culinary fruits, Berchemia lineata, Auerodendron northropianum, Berchemia scandens
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Rhamnaceae

Buckthorn family
Rhamnus pumila Turra
Scientific classification
Kingdom: Plantae
(unranked): Angiosperms
(unranked): Eudicots
(unranked): Rosids
Order: Rosales[1]
Family: Rhamnaceae
Juss.
Type genus
Rhamnus
L.
Genera

See text.

The range of Rhamnaceae.
Synonyms

Frangulaceae DC.
Phylicaceae J.Agardh
Ziziphaceae Adans. ex Post & Kuntze[2]

Rhamnaceae is a large family of flowering plants, mostly trees, shrubs, and some vines, commonly called the buckthorn family.[3]

The family contains 50–60 genera and approximately 870–900 species. Rhamnaceae have a worldwide distribution, but are more common in the subtropical and tropical regions. The earliest fossil evidence of Rhamnaceae is from the Eocene.

Leaves of Rhamnaceae family members are simple, i.e., the leaf blades are not divided into smaller leaflets.[3] Leaves can be either alternate and spiraling, or opposite. Stipules are present. These leaves are modified into spines in many genera, in some (e.g. Paliurus spina-christi and Colletia cruciata) spectacularly so. Colletia stands out by having two axillary buds instead of one, one developing into a thorn, the other one into a shoot.

Flowers of Ceanothus cuneatus
Bisexual flower of Helinus, with five sepals and petals, and a yellow, annular nectary disk. The small, clawed petals embrace the stamens.
Flowers of Ziziphus mucronata
Flowers of Rhamnus saxatilis

The flowers are radially symmetrical. There are 5 (sometimes 4) separate sepals and 5 (sometimes 4 or none) separate petals. The petals may be white, yellowish, greenish, pink or blue, and are small and inconspicuous in most genera, though in some (e.g. Ceanothus) the dense clusters of flowers are conspicuous. The 5 or 4 stamens are opposite the petals.[3] The ovary is superior, with 2 or 3 ovules (or one by abortion).

The fruits are mostly berries, fleshy drupes or nuts. Some are adapted to wind carriage, but most are dispersed by mammals and birds. Chinese jujube is the fruit of the jujube tree (Ziziphus zizyphus) and is a major fruit in China.

The American genus Ceanothus, which has several showy ornamental species, has nitrogen-fixing root nodules.

Economic uses of Rhamnaceae are chiefly as ornamental plants and as the source of many brilliant green and yellow dyes. The wood of Rhamnus was also the most favoured species to make charcoal for use in gunpowder before the development of modern propellants.

Contents

  • Genera 1
  • Systematics 2
  • References 3
  • External links 4

Genera

Tribe Ampelozizipheae
Tribe Bathiorhamneae
Tribe Colletieae
Tribe Doerpfeldieae
Tribe Gouanieae
Tribe Maesopsideae
Tribe Paliurieae
Tribe Phyliceae
Tribe Pomaderreae
Tribe Rhamneae
Tribe Ventilagineae


Incertae sedis

Systematics

Modern molecular phylogenetics recommend the following clade-based classification of Rhamnaceae:

Rhamnaceae

Ampeloziziphoids

Ventilago



Bathiorhamnus



Ampelozizyphus


Doerpfeldia




Rhamnoids

Maesopsis




Scutia



Rhamnus


Frangula





Sageretia



Berchemia




Rhamnidium


Rhamnella




Reynosia



Krugiodendron



Karwinskia


Condalia










Ziziphoids

Schistocarpeia





Hovenia



Ziziphus


Paliurus






Gouania


Helinus




Pleuranthodes



Crumenaria


Reissekia








Lasiodiscus


Colubrina




Emmenosperma




Noltea



Trichocephalus



Nesiota


Phylica







Granitites


Alphitonia


Colletieae


Adolphia


Trevoa




Discaria


Kentrothamnus


Colletia


Retanilla





Ceanothus

Pomaderreae

Siegfriedia


Cryptandra



Stenanthemum



Trymalium



Pomaderris


Spyridium














References

  1. ^ Walter S. Judd and Richard G. Olmstead (2004). "A survey of tricolpate (eudicot) phylogenetic relationships". American Journal of Botany 91 (10): 1627–1644.   (full text )
  2. ^ Juss., nom. cons."Rhamnaceae"Family: . Germplasm Resources Information Network. United States Department of Agriculture. 2003-01-17. Retrieved 2011-01-29. 
  3. ^ a b c Flowering Plants of the Santa Monica Mountains, Nancy Dale, 2nd Ed. 2000, p. 166
  4. ^ "Granitites".  
  5. ^ "Rhamnaceae"GRIN Genera of . Germplasm Resources Information Network. United. Retrieved 2011-01-29. 

External links

  • Rhamnaceae of Mongolia in FloraGREIF
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